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Abstract During the summer when an El Niño event is transitioning to a La Niña event, the extratropical teleconnections exert robust warming anomalies over the U.S. Midwest threatening agricultural production. This study assesses the performance of current climate models in capturing the prominent observed extratropical responses over North America during the transitioning La Niña summer, based on atmospheric general circulation model experiments and coupled models from the North American Multimodel Ensemble (NMME). The ensemble mean of the SST-forced experiments across the transitioning La Niña summers does not capture the robust warming in the Midwest. The SST-forced experiments do not produce consistent subtropical western Pacific (WP) negative precipitation anomalies and this leads to the poor simulations of extratropical teleconnections over North America. In the NMME models, with active air–sea interaction, the negative WP precipitation anomalies show better agreement across the models and with observations. However, the downstream wave train pattern and the resulting extratropical responses over North America exhibit large disagreement across the models and are consistently weaker than in observations. Furthermore, in these climate models, an anomalous anticyclone does not robustly translate into a warm anomaly over the Midwest, in disagreement with observations. This work suggests that, during the El Niño to La Niña transitioning summer, active air–sea interaction is important in simulating tropical precipitation over the WP. Nevertheless, skillful representations of the Rossby wave propagation and land–atmosphere processes in climate models are also essential for skillful simulations of extratropical responses over North America.